A brief note on Secondary Growth: how the vascular cambium and the cork cambium make a dicot stem thicker, annual rings, heartwood and sapwood, and the bark and lenticels, with a worked example and questions.
A brief note on Secondary Growth. Read it before a test. Try each question at the end before you open its answer.
Growing thicker
Secondary growth makes dicot stems and roots thicker. Lateral meristems do it. Monocots have closed bundles and no cambium, so they show none.
Vascular cambium: fascicular cambium in the bundles plus interfascicular cambium from the medullary rays. Together they form a ring.
It cuts secondary xylem inward and secondary phloem outward. It makes much more xylem. Wood is secondary xylem.
Ray initials make secondary medullary rays.
The outer cover
Cork cambium (phellogen) arises in the cortex. It cuts cork (phellem) outward and phelloderm inward. The three make the periderm.
Cork is dead, with suberin in its walls. It is waterproof and airtight.
Lenticels are patches of loose cells in the cork. They let gases pass.
Bark means all the tissues outside the vascular cambium.
Rings and wood
Spring wood: wide vessels, light, low density. Autumn wood: narrow vessels, dark, high density. One of each makes an annual ring.
The number of rings gives the age. Wide rings mean good years.
Heartwood: dead, dark with tannins and resins, hard, does not conduct. Sapwood: outer, light, carries water and minerals.
A worked example
A tree is felled and its stump shows a pale, soft outer ring of wood and a dark, hard inner core. A hollow in the core has no effect on the leaves above.
A dark, hard core. It is heartwood, the oldest wood. It is dead.
A pale outer ring. It is sapwood, which carries the water.
Leaves unaffected by the hollow. Water still moves up the sapwood, so the tree lives.
Practice with new numbers
A program to try
The program lists the layers of a woody stem from the outside inwards, with the job of each.
Mistakes to avoid
Giving monocots secondary growth. Their bundles are closed and have no cambium.
Saying the cambium cuts xylem outward. Xylem is cut inward and phloem is cut outward.
Saying heartwood carries water. It is dead. Sapwood carries water.
Counting two rings for each year. Spring wood and autumn wood together make one ring.